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Medicine 1902

The Mechanism of Pancreatic Secretion

William Bayliss & Ernest Starling

Acid in the gut sent a chemical through the blood to the pancreas — the first hormone, and the birth of endocrinology.

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In depth · the introduction

The body, everyone assumed, was wired like a telephone network. Two scientists in London found it also runs a postal service — and posted the first letter.

The big idea

Bayliss and Starling discovered that organs talk to each other not only through nerves, but through chemicals released into the blood. When acid arrives in the gut, the gut lining drops a chemical into the bloodstream; the blood carries it to the pancreas, which then pours out the juice that neutralises the acid. They called the chemical secretin.

Three years later Starling gave a name to this whole new class of blood-borne messengers: hormones. Every hormone you've heard of — adrenaline, insulin, testosterone — is the same kind of signal. This was the first one ever caught in the act, and it began the science of endocrinology.

How it came about

In 1902, the famous Russian physiologist Ivan Pavlov had convinced the world that digestion was run by nerves — reflexes triggered by food and acid. Bayliss and Starling, working together at University College London (and married into the same family — Bayliss had married Starling's sister), decided to test whether the gut-to-pancreas signal really needed nerves.

They took a loop of a dog's intestine and cut every nerve to it, leaving only its blood supply. Then they dripped acid inside. The pancreas, far away, secreted anyway. The signal couldn't be travelling by nerve — it had to be in the blood. To prove it, they scraped the gut's lining, soaked it in acid, and injected the liquid into a vein: the pancreas gushed. So thrilled were they that they reported it to the Royal Society just a week later.

Why it mattered

It revealed a second control system for the whole body. Nerves are like private phone lines — fast and aimed at one target. Hormones are like a message poured into a river: they reach everyone, but only the organ tuned to that message acts on it. Understanding hormones is what later let doctors treat diabetes with insulin, an underactive thyroid with thyroxine, and much more. It is one of the foundations of modern medicine.

A way to picture it

Imagine the gut needs to tell the pancreas, "acid's coming — send help." A nerve would be a phone call: a wire from one to the other. But Bayliss and Starling showed the gut instead tosses a sealed note into the bloodstream, like a message in a bottle dropped in a river. The note floats past every organ, but only the pancreas can read it — and when it does, it acts. Cut the phone lines and the note still arrives, because it was never travelling by wire.

An interactive schematic: a gut and a pancreas joined by a blood path and a severable nerve path, above a dose-response curve. A slider raises the acid stimulus and the pancreas's secretion climbs a saturating curve; a button cuts the nerves, marking them with a cross, yet the secretion is unchanged — the signal travels by blood.

Where it sits

This was a turning point between two giants' worldviews. Pavlov — also in this Library, for his work on conditioned reflexes — had made nerves the master of digestion; Bayliss and Starling showed chemistry shared the throne. The hormone idea they launched runs straight on to Banting and Best's insulin two decades later, to the discovery of adrenaline and the sex hormones, and to today's gut-hormone drugs like the GLP-1 agonists used for diabetes and weight loss. Endocrinology begins here.

The original document
Original source text
W. M. Bayliss & E. H. Starling · J. Physiol. 28 (1902): 325–353
The question: nervous or chemical?
The paper opens on a known fact — that acid entering the small intestine provokes a flow of pancreatic juice — and on the prevailing explanation, descended from Pavlov's school, that this was a nervous reflex carried along the nerves of the gut.
The decisive experiment
Bayliss and Starling isolated a loop of jejunum in an anaesthetised dog and cut every nerve to it, leaving it connected only by its blood vessels. They introduced dilute acid into the loop — and the pancreas, at a distance, still poured out juice. A nervous reflex was ruled out: the signal had to travel by blood.
Secretin
They then scraped the lining of the jejunum, ground it with sand and dilute acid, filtered the result, and injected the filtrate into a vein. The pancreas responded as vigorously as to acid in the gut itself. The active substance — released from the gut wall into the blood by acid — they named secretin.
[ … ]
Conclusion
The authors conclude that pancreatic secretion is governed not by a nerve but by a chemical messenger produced in the intestinal lining and carried in the bloodstream. The full paper — with its seventeen figures and secretion tracings — is available at the source below.
University College London · 1902